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This is especially likely to happen when the cast of a series functions as a Universal-Adaptor Cast.
is a Japanese term for a character development process that describes a person who is initially cold and even hostile towards another person before gradually showing a warmer side over time.
Comiket organizer Koichi Ichikawa has described Lum Invader of Urusei Yatsura as being both the source of moe and the first tsundere; Tsundere as a concept is not strictly limited to women, and is not strictly limited to manga or anime.
The character Germany from the series Hetalia: Axis Powers is portrayed as being tsundere, and is paired with a "lovable loser", Italy Veneziano.
Instrumentality is a key theme throughout the series, and some say that, in the end of the TV series, Shinji finally accepts instrumentality.